This work was supported by the National Natural Science Foundation of China (grant no. 31872877 and grant no. 32071940), the Key R&D projects in Hunan Province (grant no. 2019NK206102 and grant no. 2020NK2028), and the special project for post experts of the modern agricultural industrial technology system in Hunan Province (2019–2023), National Key R&D projects (grant no. 2019YFD1002205-3).