Conceptualization, L.L. and Z.L.; methodology, D.L.; software, Z.L. and N.L.; validation, L.L. and Z.L.; formal analysis, N.Z.; investigation, N.L., K.T., X.L., Y.T. and X.Y.; resources, L.L.; data curation, N.L., K.T., X.L., Y.T. and X.Y.; writing—original draft preparation, N.L.; writing—review and editing, N.L., L.L. and Z.L.; visualization, N.L.; supervision, D.L., N.Z., L.L. and Z.L.; project administration, D.L., N.Z., L.L. and Z.L.; funding acquisition, D.L., N.Z., L.L. and Z.L. All authors have read and agreed to the published version of the manuscript. This work was supported by the National Key R&D Project (No. 2018YFD1000900), Chinese Agricultural Research System (CARS-13), Hunan Key R&D Project (No. 2021NK2005), Shandong Key R&D Project (No. ZFJH202310-2), and Hunan Modern Agricultural Research System. This work was also funded by General Scientific Research Project of Education Department of Hunan Province (19C0930) and Crop Science Excellent Talent Fund of Hunan Agricultural University (YXQN2018-21).