Conceptualization, M.B. and H.L.; Data curation, S.W. and M.B.; Formal analysis, S.W.; Funding acquisition, R.H. and H.L.; Investigation, Y.S. and Z.Y.; Supervision, K.X., X.X., R.H., Y.L. and H.L.; Validation, Y.L.; Writing—original draft, S.W.; Writing—review & editing, M.B. and H.L. All authors have read and agreed to the published version of the manuscript. The project was supported by grant from was supported by Science and technology program of Changsha (kq1907074), grant from National Natural Science Foundation of China (32072741), and the Innovation Team in Key Area Innovation Team of Physiology and Metabolism and Body Health in Pig (2019RS3022). The research was also funded by the research program of Huxiang Youth Talent Program of Hunan Province (2018RS3110), Youth Innovation Promotion Association, CAS (2019356), and the Special Funds for the Construction of Innovative Provinces in Hunan (2019RS1068, 2019NK2193).