Acknowledgements The study was financially supported by National Natural Science Foundation of China (NSFC; Grant Nos. 51709103, 51409024, 51222805, 51408219, 51579096, 51521006, and 51508175), The Special Environmental Protection Foundation for Public Welfare Project (201509032), Project of Science and Technology of Hunan Province (2015WK3016), and Project of Science and Technology of Hunan Province (2016WK2010).