This work was supported by Joint Funds of National Natural Science Foundation of China [grant number U19A2030], the National Natural Science Foundation of China [grant number 31271789], Central Committee Guides Local Science and Technology Development Program [grant number 2019XF5041], and the Special Project for the Construction of Modern Agricultural Industrial Technology System in Hunan Province [Xiangcai Nongzhi; grant number 20190047]. CRediT authorship contribution statement Chenyu Zhang: Conceptualization, Investigation, Writing - original draft. Xiaoqin Yi: Writing - review & editing, Supervision. Fang Zhou: Resources, Formal analysis. Xizhi Gao: Data curation. Minhan Wang: Software. Jianjiao Chen: Validation. Jianan Huang: Project administration. Chengwen Shen: acquisition.