Conceptualization, S.Z., W.W. and L.L.; Data curation and formal analysis, S.Z., C.L., J.W. and S.P.; Funding acquisition, H.M., W.W. and L.L.; Investigation and methodology, S.Z. and C.L.; Project administration and resources, W.W. and L.L.; Software, S.Z., C.L., J.W. and S.P.; Supervision and validation, W.W. and L.L.; Visualization, S.Z. and C.L.; Writing—original draft, S.Z.; Writing—review & editing, L.L. All authors have read and agreed to the published version of the manuscript. This study had the support of the Department of Agriculture and Rural Affairs of Hunan Province; we also acknowledge financial support from Department of Agriculture and Rural Affairs of Hunan Province. The authors also acknowledge the support from the Scientific Research Innovation Program of Hunan (CX20190507), Natural Science Foundation of Hunan Province (2019JJ50262), and the Hunan Engineering Technology Research Center for Rapeseed Oil Nutrition Health and Deep Development.