This study was supported by the National Natural Science Foundation of China (Nos. 31772642, 31672457), the Ministry of Agricultural of the People’s Republic of China (2015-Z64, 2016-X47), the Hunan Provincial Science and Technology Department (2021JJ30008, 2019TP2004, 2017NK2322, 2016WK2008, 2016TP2005), the Double first-class construction project of Hunan Agricultural University (SYL201802003), the China Postdoctoral Science Foundation (2018M632963, 2019T120705), the Postgraduate Scientific Research Innovation Project of Hunan Province (CX20210654), and the Science and Technology Innovation and Entrepreneurship Project for University Students of Hunan Province (2021RC1004).