The author(s) declare financial support was received for the research, authorship, and/or publication of this article. Funding for this research was provided by the National Key R&D Program of China, 2022YFD1401200; Central Public-interest Scientific Institution Basal Research Fund (No. IVF-BRF2022013), Key Laboratory of Biology and Genetic Improvement of Horticultural Crops, Ministry of Agriculture, P.R. China (CARS-25), the Science and Technology Innovation Program of the Chinese Academy of Agricultural Sciences (CAAS-ASTIP-2017-IVF), the National Natural Science Foundation of China (grant number: 32372508); The Forestry Science and Technology Innovation Foundation of Hunan Province for Distinguished Young Scholarship (XLKJ202205), the Foundation of Changsha Municipal Science and Technology Bureau (KQ2202227), the key project of Hunan Provincial Education Department (22A0155), the Forestry Bureau for Industrialization Management of Hunan Province (2130221), and the Graduate Innovation Project of Hunan Province (2023XC108), the College Student Innovation and Entrepreneurship Project of China (S202310537005). The Project fund of Hunan Province Philosophy and social science achievements Evaluation Committee (XSP20YBZ123).