This study was financially supported by the National Natural Science Foundation of China [Grant No. 21407182 and 51709103 ], Natural Science Foundation of Hunan Province of China [Grant No. 2018jj2490 ], Special Fund for Agro-scientific Research in the Public Interest of China [Grant No. 201503108 ], National Key R&D Projects of China [Grant No. 2016YFC0403002 ], Science & Technology Innovation Platform and Talent Project of Hunan Province of China [Grant No. 2018WK4007 ] and Double first-class construction project of Hunan Agricultural University [Grant No. SYL201802005 ].