This research was supported by China Agriculture Research System of MOF and MARA (CARS-29). CRediT authorship contribution statement Ning Shi: Writing – original draft, Visualization, Investigation, Formal analysis. Xuan-Xuan Pei: Investigation. Ming-Yu Li: Investigation. Wen-Ting Chen: Investigation. Hui-Qing Li: Investigation. Guo-Shun Yang: Supervision. Chang-Qing Duan: Supervision. Jun Wang: Writing – review & editing, Supervision, Project administration, acquisition, Conceptualization.