The present work was financially supported by the National Natural Science Foundation of China (Grant No. 32302764), Hunan Provincial Natural Science Foundation (Grant No. 2024JJ5179), Hunan Provincial Education Department (Grant No. 22A0168), Science and Technology Department of Xinjiang Uygur Autonomous Region (Grant No. 2022A02001-1), Hunan Provincial Science and Technology Department (Grant No. 2021RC4060), and Hunan Herbivores Industry Technological System.