This research was funded by the National Key Research and Development Program of China, grant number 2021YFD1600100, the Fundamental Research Program of Shanxi Province (grant number: 202203021222178), and the Excellent Doctoral Program Initiated by Shanxi Agricultural University (2023BQ32). CRediT authorship contribution statement Wang Haizhen: Methodology, Investigation. Zhao Mengying: Writing – review & editing, Writing – original draft, Methodology, Investigation, Conceptualization. Yang Dan: Investigation. Li Zhihao: Methodology, Investigation. Wang Han: Investigation. Liu Zhenyu: Writing – review & editing. Fu Yongxia: Writing – review & editing, Supervision, Project administration, Investigation. Guo Shang: Writing – review & editing, Supervision, Project administration, acquisition, Conceptualization.