Conceptualization, X.Y. and W.L.; methodology, X.Y. and D.Z.; software, X.Y. and D.Z.; validation, X.Y., W.L., L.Q. and D.Z.; formal analysis, X.Y.; investigation, X.Y., D.Z. and W.L.; resources, X.Y., W.L., L.Q. and D.Z.; data curation, X.Y., W.L., L.Q. and D.Z.; writing—original draft preparation, X.Y.; writing—review and editing, W.L. and L.Q.; visualization, X.Y.; supervision, W.L. and L.Q.; project administration, W.L.; funding acquisition, W.L. All authors have read and agreed to the published version of the manuscript. This research was funded by the Hunan Social Science Fund Project (19YBA195) and the Postgraduate research and innovation project of Hunan Agricultural University (2022XC043).