Conceptualization, J.Z.; Data curation, J.Z.; Formal analysis, J.Z. and H.H.; Funding acquisition, L.Q. and Y.L.; Investigation, J.Z., L.Q., Q.L., Y.Z., J.S., Q.G., H.H., W.D. and J.X.; Methodology, J.Z.; Project administration, Y.L.; Resources, J.S. and Y.L.; Software, J.Z. and Q.G.; Supervision, L.Q., W.D. and Y.L.; Validation, J.Z., Q.L. and Y.Z.; Writing—original draft, J.Z.; Writing—review and editing, L.Q. and Y.L. All authors have read and agreed to the published version of the manuscript. This work was supported by the Science and Technology Innovation Program of Hunan Province (2022RC1025), the National Natural Science Foundation of China (Grant No. 31801736) and the Double first-class construction project of Hunan Agricultural University (Grant No. SYL2019029).