Conceptualization, S.Q.; methodology, X.F. and Y.T.; software, X.F. and Y.T.; validation, M.S., X.F. and Y.T.; formal analysis, Y.T.; investigation, C.Z.; resources, S.Q.; data curation, X.F. and Y.T.; writing—original draft preparation, X.F.; writing—review and editing, X.F.; visualization, M.S., X.F. and Y.T.; supervision, S.Q.; project administration, M.S.; funding acquisition, S.Q. All authors have read and agreed to the published version of the manuscript. This research was funded by the National Key Research and Development Program of China (2019YFC1604903), Natural Science Foundation of China (31101268), Natural Science Foundation of Hunan Province (2019JJ40132), and Double First-Class Construction Project of Hunan Agricultural University (SYL201802025) to Si Qin.