Conceptualization, X.L. and Z.D.; methodology, P.C., J.L. and Q.X.; software, Q.X. and L.C.; validation, X.L., Z.D., Q.T. and L.C.; formal analysis, J.L., T.Z., Y.X. and L.G.; investigation, P.C. and J.L.; resources, J.L. and P.C.; data curation, P.C., T.Z. and L.G.; writing—original draft preparation, P.C., Q.T. and L.G.; writing—review and editing, P.C. and J.L.; visualization, T.Z., Y.X. and L.G.; supervision, X.L. and Z.D.; project administration, X.L.; funding acquisition, X.L. and P.C. All authors have read and agreed to the published version of the manuscript. This research was funded by the Agricultural Science and Technology Innovation Funds Project of Hunan Province (2023CX71, 2023CX23), the National Natural Science Foundation of China (32302487), the Natural Science Foundation of Changsha City (kq2208136), and the National Key Research and Development Program of China (2022YFF1003100, 2019YFD1001400).