Conceptualization, H.Q., Y.S. and Y.H.; Methodology, H.Q., C.L., Q.S., S.D., X.D., L.C., X.C., Y.S. and Y.H.; Software, H.Q., Q.S., S.D., X.D. and L.C.; Formal analysis, H.Q., C.L., X.C. and Y.H.; Investigation, C.D., L.C. and Y.H.; Writing—original draft, H.Q. and C.L.; Writing—review & editing, H.Q., C.L. and Y.H.; Visualization, H.Q.; Supervision, Y.H.; Project administration, X.C., Y.S. and Y.H.; Funding acquisition, X.C., Y.S. and Y.H. All authors have read and agreed to the published version of the manuscript. This study was financially supported by the National Key Research Program (2017YFC0505503) and the National Natural Science Foundation of China (42177295, 42277309).