This study was partially supported by the Shenzhen Strategic Emerging Industry Development Special Fund Project (Shenzhen Municipal Economic and Trade Information Commission, #20180130150742764 to SL), United States Department of Agriculture (MAS00450 and MAS00492, and NIFA grant #2019-67017-29249 and 2020-67017-30835 to HX), Hunan Science and Technology Plan Program (2019RS1055 to ZT), and Major Research plan of Changsha (kq1801016 to ZT).