Conceptualization, Y.Z. and L.L.; methodology, D.X., B.W., H.L. (Hong Lu), T.P., J.S. and H.L. (Hongyu Lei); validation, Y.Z. and L.L.; formal analysis, L.L. and J.H.; investigation, D.X., B.W., H.L (Hong Lu). and T.P.; data curation, D.X., J.H. and L.L.; writing—original draft preparation, L.L.; writing—review and editing, J.S. and H.L. (Hongyu Lei); visualization, D.X. and L.L.; supervision, L.L.; project administration, Y.Z. and L.L.; funding acquisition, L.L. All authors have read and agreed to the published version of the manuscript. This research was funded by the Science Fund for Distinguished Young Scholars of Changsha City, grant number kq2009055, and the Scientific Research Foundation for the introduction of talent in Hunan Agricultural University, grant number 15YJ05.