National Natural Science Foundation of China [51974123]; Distinguished Youth Foundation of Hunan Province [2020JJ2018]; Key R & D projects in Hunan Province [2020WK2016, 2020SK2032, 2021SK2047, 2022NK2044]; Natural Science Foundation of Hunan Province, China [2021JJ40261]; Hunan High Level Talent Gathering Project [2019RS1077, 2020RC5007]; Natural Sciences Foundation of Hunan Agricultural University, China [19QN11]; Hunan Provincial Key Laboratory of Crop Germplasm Innovation and Resource Utilization Science Foundation [19KFXM12]; Changsha Science and technology plan [KH2005114, KH2201428]; Hunan Graduate Scientific Research Innovation Project [2022XC017]