Conceptualization, S.C., J.T. and Z.L.; Data curation, Z.H., Y.C., Y.X., G.Z., Y.L. and J.L.; Formal analysis, X.L., F.Q. and S.L.; Funding acquisition, Z.H. and Z.L.; Investigation, Y.C., Y.X., G.Z., X.L., Y.L. and J.L.; Methodology, Z.H., Y.C., S.C., F.Q., J.T. and S.L.; Project administration, Z.L.; Resources, Z.L.; Supervision, S.L.; Writing—original draft, Z.H. and Y.C.; Writing—review and editing, Z.H., Z.L. and S.L. All authors have read and agreed to the published version of the manuscript. This work was funded by National Natural Science Foundation of China (Grant No. U19A2041, 31902345 and 321102813); Hunan Provincial Natural Science Foundation of China (Grant No. 2023JJ30075 and 2021JJ40627); Key program of Hunan education committee (Grant No. 22A0597).