Conceptualization, Y.H. and C.L.; methodology, C.L., Y.H. and Z.X.; software, Y.H.; validation, W.Z. (Wan Zhang) and J.H.; formal analysis, Y.H.; investigation, C.L.; resources, Y.Z. and Z.X.; data curation, Z.X.; writing—original draft preparation, Y.H.; writing—review and editing, W.Z. (Wan Zhang) and Z.X.; visualization, Y.H., C.L. and Z.X.; supervision, Z.S., W.Z. (Weiping Zhao); project administration, Y.Z. and Z.X.; funding acquisition, Y.Z. and Z.X. All authors have read and agreed to the published version of the manuscript. This research was funded by the Key Projects of National Forestry and Grassland Bureau (201801), China Postdoctoral Science Foundation (2020M683592), and Open Fund of Key Laboratory of Microbial Resources Collection and Preservation, Ministry of Agriculture and Rural Affairs (KLMRCP2021-07). The funding bodies played no role in the design of the study and collection, analysis, and interpretation of data and in writing the manuscript.