Conceptualization, B.Z. and Y.X.; Methodology, Y.L., L.T., B.Z. and Y.X.; Validation, Y.L., L.T.; Formal analysis, Y.L.; Investigation, L.T., G.H., L.L., C.D., H.L. and B.M.; Data curation, Y.L., G.H., L.L., C.D. and H.L.; Writing—original draft, Y.L.; Writing—review & editing, L.T., B.M., B.Z. and Y.X.; Visualization, Y.L.; Project administration, L.T.; Funding acquisition, Y.L., L.T. and B.Z. All authors have read and agreed to the published version of the manuscript. This research was supported by the National Natural Science Foundation of China (32101681), the Natural Science Foundation of Hunan Province (2022JJ30034), the Earmarked Fund for China Agriculture Research System (CARS-01-14), the Pilot Project for Remediation and Management of Heavy Metal-Contaminated Farmland and Adjustment of Crop Planting Structure in the Changsha-Zhuzhou-Xiangtan region (XCNZ [2018] 83, [2019] 33, [2020] 28), and the Genetically Modified Organisms Breeding Major Projects (2016ZX08001004).