This study was supported by the Major Basic Research Projects of Shandong Natural Science Foundation (ZR2018ZC2363) and Major Science and Technology Innovation Projects in Shandong Province (2021CXGC010804). Author contributions Zerun Yin, Investigation, Writing – original draft; Jinpeng Yu, Investigation, Resources data curation; Xinran Han, Data curation; Hui Wang, Software; Quangang Yang, Formal analysis; Hong Pan, Methodology; Yanhong Lou, Conceptualization, acquisition, Writing – review & editing; Yuping Zhuge, Conceptualization, acquisition, Project administration.