Conceptualization, Y.Z. and Y.W.; methodology, Y.Z., T.L. and Y.W.; software, Y.Z.; validation, Y.Z. and Y.W.; formal analysis, Y.Z., L.C., X.Z. and T.X.; investigation, Y.Z., L.C., X.Z. and T.X.; data curation, Y.Z., T.L. and Y.W.; writing—original draft preparation, Y.Z.; writing—review and editing, Q.S., T.L. and Y.W.; supervision, T.L.; project administration, Y.W.; funding acquisition, T.L. and Y.W. All authors have read and agreed to the published version of the manuscript. This study was supported by the National Natural Science Foundation of China (32072334) and the National Natural Science Foundation of Hunan Province, China (2020JJ4362), Agricultural Science and Technology Innovation Fund project of Hunan Province (2022CX02).