This study was supported by grants from the National Natural Science Foundation of China (31801721; 31672017), National Key Research and Development Project (2016YFD0200800; 2016YFD0300707), the Natural Science Foundation of Hunan Province, China (2020JJ5240), the Scientific Research Fund of Hunan Provincial Education Department (19B247), Commonweal Plan Project of Ningbo City (202002N3004), and the Youth Fund Project of Hunan Agricultural University (19QN31).