This work was financially supported by the National Natural Science Foundation of China ( 21878326 ), the Science and Technology Innovation Program of Hunan Province-Huxiang Young Talent Program ( 2021RC3116 ), the Training Program for Excellent Young Innovators of Changsha ( kq2106073 ), the Agricultural Science and Technology Innovation Program ( ASTIP-IBFC08 ), and the earmarked fund for the China Agriculture Research System ( CARS-16-E24 ).