R.Z.: Investigation, data curation, visualization, writing—original draft preparation; H.L.: supervision, conceptualization, methodology, investigation, data curation; R.Z., M.D. and D.L.: resources, formal analysis, software; H.L., Y.L., M.D. and B.T.: resources, project administration, conceptualization, writing—review and editing; H.L.: funding acquisition, supervision, conceptualization, visualization, writing—review and editing. All authors have read and agreed to the published version of the manuscript. This research was funded by grants from the Crop Varietal Improvement and Insect Pests Control by Nuclear Radiation, the Key R&D Plan of Hunan Province (2022NK2002).