National Key R&D Program of China [2017YFD0200100, 2017YFD0200103]; National Natural Science Foundation of ChinaNational Natural Science Foundation of China (NSFC) [31101596, 31372130]; Hunan Provincial Recruitment Program of Foreign Experts; National Oilseed Rape Production Technology System of China; "2011 Plan" - Ministry of Education of ChinaMinistry of Education, China