Conceptualization, X.Z. and Y.H.; methodology, J.Z. (Jing Zhang), X.H. and J.Z. (Jun Zhou); software, L.Y., S.P. and H.Y.; validation, L.Y. and S.P.; formal analysis, H.Y., Z.D., S.P. and Q.T.; investigation, X.H., J.Z. (Jun Zhou), L.Y. and X.Z.; resources, X.Z. and Y.H.; data curation, J.Z. (Jing Zhang), X.H., J.Z. (Jun Zhou) and Z.D.; writing—original draft preparation, J.Z. (Jing Zhang) and X.H.; writing—review and editing, J.Z. (Jing Zhang), Q.T. and Y.H.; visualization, J.Z. (Jing Zhang) and Y.H.; supervision, Y.H.; project administration, Q.T. and X.Z.; funding acquisition, J.Z. (Jing Zhang) and Y.H. All authors have read and agreed to the published version of the manuscript. This work was funded by the China Postdoctoral Science Foundation, grant number 2023M741143, the Scientific Research Fund of Hunan Provincial Education Department, grant number 23B0228, and the Research Innovation Project of Postgraduates in Hunan Agricultural University, grant number CX20230706.