Our work has been fully supported by the Key Research and Development Program of Hunan Province, China (Grant No. 2017NK2381), the Double First-class Construction Project of Hunan Agricultural University, China (Grant No. SYL2019077), the National Natural Science Foundation of China (Grant No. 61972146) as well as the Natural Science Foundation of Hunan Province, China (Grant No. 2019JJ40133).