This study was financially supported in part by the National Key R&D Program of China (2017YFD0200100), National Natural Science Foundation of China (Grant No. 31101596, 31372130), Hunan Provincial Recruitment Program of Foreign Experts, the National Oilseed Rape Production Technology System of China; “2011 Plan” supported by The Chinese Ministry of Education, and Research and Innovation Project of Postgraduates in Hunan Province (CX2015B242).