Conceptualization, C.-F.L.; Methodology, Y.-J.D.; Validation, Y.-J.D. and J.-L.L.; Formal analysis, X.-H.Z.; Investigation, C.-F.L.; Data curation, Z.-D.W. and T.H.; Writing-original draft, C.-F.L. and Z.-D.W.; Writing-review & editing, Y.-J.D. and J.-L.L.; Visualization, T.H.; Funding acquisition, X.-H.Z. All authors have read and agreed to the published version of the manuscript. This research was funded by the Hunan Provincial Key Research and Development Program under Grant 2023NK2011 and the Meizhou Tobacco Science Research Project under Grant No. 202204.