Conceptualization, L.X., X.B. and H.Z.; methodology, N.Z. and X.Z.; validation, P.Y., H.H. and J.L.; investigation, N.Z. and P.Y.; resources, X.Z. and J.L.; data curation, N.Z. and X.Z.; writing—original draft preparation, N.Z.; writing—review and editing, X.Z. and H.H.; supervision, L.X.; project administration, L.X., X.B. and H.Z.; funding acquisition, L.X., X.B. and H.Z. All authors have read and agreed to the published version of the manuscript. This research was funded by the National Tea Industry Technology System Project, grant number CARS–19; the Key Research and Development Project of Hunan Province, grant number 2021NK1020–2; and the Agricultural Science and Technology Innovation Funding Program of Hunan Province, grant number 2020CX34.