Qian Lin is a student of the Joint PhD Program supported by China Scholarship Council. This research was supported by the Special Fund for National Key R & D Program (Grant: 2016YFD0501209), Agro-scientific Research in the Public Interest (Grant: 201403047), National Science and Technology Basic Project (Grant: 2014FY111000-3), Hunan Key Scientific and Technological Project (Grant: 2016NK2124), Core Research Program 1515 of Hunan Agricultural University, Hunan Postgraduate Scientific Research and Innovation Projects (Grant: CX2014B284), Excellent Doctoral Dissertation Cultivating Fund of Hunan Agricultural University (Grant: YB2015003).