Conceptualization, X.W. and S.Q.; methodology, X.W., C.L. and S.Q.; software, X.W. and C.L.; validation, C.L. and R.Z.; formal analysis, X.W.; investigation, X.W., C.L. and Y.W.; resources, S.Q.; data curation, C.L. and Y.W.; writing—original draft preparation, X.W. and C.L.; writing—review and editing, S.Q.; visualization, X.W. and S.Q.; supervision, X.Z. and S.Q.; project administration, X.Z. and S.Q.; funding acquisition, S.Q. and X.Z. All authors have read and agreed to the published version of the manuscript. This research was funded by the National Key Research and Development Program of China (2019YFC1604903) and Natural Science Foundation of Hunan Province (2023JJ30295) to Si Qin; and Natural Science Foundation of Changsha City (kq2208076) to Xixin Zhou.