CRediT authorship contribution statement Zemao Yang: Conceptualization, Data curation, Formal analysis, acquisition, Investigation, Methodology, Project administration, Resources, Software, Visualization, Writing – original draft, Writing – review & editing. Canhui Deng: Data curation, Investigation, Resources, Software, Validation. Yupeng Wu: Data curation, Investigation, Methodology, Software, Validation, Visualization. Zhigang Dai: Investigation, Resources. Qing Tang: Formal analysis. Chaohua Cheng: Formal analysis. Ying Xu: Acknowledgments This study was financially supported by the National Natural Science Foundation of China (grant number 31601351), the Science and Technology Innovation Project of Chinese Academy of Agricultural Sciences (grant numbers CAAS-ASTIP-2017-IBFC01 and CAAS-XTCX2016016-2), National Key Research and Development Project (2017YFD0101703), and Major National Genetically Modified Project (SQ2018ZD080068). The funding bodies did not participate in the design of the study, the collection, analysis, and